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the protein, and reading of the messenger RNA continues. At least three suppressors of the mutant amber codon (UAG) are known in E. coli. One suppressor puts tyrosine, one puts glutamine, and one puts serine into the protein chain at the point of an amber codon. Normally, tyrosine transfer RNA has the anticodon 3 -AUG5 . The suppressor transfer RNA that reads amber as a tyrosine codon has the anticodon 3 -AUC-5 , which is complementary to amber. Hence, a mutated tyrosine transfer RNA reads amber as a tyrosine codon (fig. 12.26b). If the amber nonsense codon is no longer read as a stop signal, then won t all the genes terminating in the amber codon continue to be translated beyond their ends, causing the cell to die In the tyrosine case, two genes for tyrosine transfer RNA were found; one contributes the major fraction of the transfer RNAs, and the other, the minor fraction. It is the minor-fraction gene that mutates to act as the suppressor. Thus, most messenger RNAs are translated normally, and most amber mutations result in premature termination, although a suf cient number are translated (suppressed) to ensure the viability of the mutant cell. In general, intergenic suppressor mutants would be eliminated quickly in nature because they are inefficient the cells are not healthy. In the laboratory, we can provide special conditions that allow them to be grown and studied.

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around the equivalent gate circuit of Figure 933(b) yields: VGG = vGSQ + iGQ RG + iDQ RS = vGSQ + iDQ RS where VGG = VDD /2 and RG = R1 R2 Note that iGQ = 0 because of the in nite input resistance of the MOSFET

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p-Channel MOSFETs and CMOS Devices As the designation indicates, a p-channel MOSFET is characterized by p-type doping; the construction and symbol are shown in Figure 934 Note the opposite direction of the arrow to indicate that the pn junction formed by the channel and substrate is now in the opposite direction The direction of drain current is opposite; therefore vDS and vGS are now negative A more convenient reference is obtained if voltages are de ned in the direction opposite to that for the n-channel device: if one de nes vSD = vDS and vSG = vGS , then these voltages will be positive for the drain current direction indicated in Figure 934 The carriers are holes in this device, since the channel, when formed, is p-type Aside from the nature of the charge carriers and the direction of current and polarity of the voltages, the pchannel and n-channel transistors behave in conceptually the same way; however, since holes are in general less mobile (recall the discussion of carrier mobility in Section 81), p-channel MOSFETs are not used very much by themselves They do nd widespread application in complementary metal-oxide-semiconductor (CMOS) devices CMOS devices take advantage of the complementary symmetry of p- and n-channel transistors built on the same integrated circuit Focus on Measurements: MOSFET Bidirectional Analog Gate illustrates one application of CMOS technology

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Whereas intergenic suppressors represent mutations that restore the normal phenotype, mostly through mutation of transfer RNA loci, mutator and antimutator mutations cause an increase or decrease in the overall mutation rate of the cell. They are frequently mutations of DNA polymerase, which, as you remember, not only polymerizes DNA nucleotides 5 3 complementary to the template strand, but also checks to be sure that the correct base was put in (they proofread). If, in the proofreading process, the polymerase discovers an error, it can correct this error with its 3 5 exonuclease activity. Mutator and antimutator mutations sometimes involve changes in the polymerase s proofreading ability (exonuclease activity). Phage T4 has its own DNA polymerase with known mutator and antimutator mutants. Mutator mutants are very poor proofreaders (they have low exonuclease-topolymerase ratios), and thus they introduce mutations throughout the phage genome. Antimutator mutants, however, have exceptionally ef cient proofreading ability (high exonuclease-to-polymerase ratios) and, therefore, result in a very low mutation rate for the whole genome (box 12.3).

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convert tiff to pdf c# itextsharp

convert tiff to pdf c# itextsharp

convert tiff to pdf c# itextsharp

convert tiff to pdf c# itextsharp

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convert tiff to pdf c# itextsharp

convert tiff to pdf c# itextsharp

convert tiff to pdf c# itextsharp

convert tiff to pdf c# itextsharp